Neal E. A. Kroll

Mild hypoxia disrupts recollection, not familiarity

Authors:

  • Andrew P. Yonelinas

  • Joel R. Quamme

  • Keith F. Widaman

  • Neal E. A. Kroll

  • Mary Jane Sauve

  • Robert T. Knight

Date: 2004

PubMed: 15535174

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Abstract:

Yonelinas et al. (2002) found that hypoxic patients exhibited deficits in recollection that left familiarity relatively unaffected. In contrast, Manns, Hopkins, Reed, Kitchener, and Squire (2003) studied a group of hypoxic patients who suffered severe and equivalent deficits in recollection and familiarity. We reexamine those studies and argue that the discrepancy in results is likely due to differences in the hypoxic groups that were tested (i.e., differences in amnestic severity, subject sampling methods, and patient etiology). Yonelinas et al. examined memory in 56 cardiac arrest patients who suffered a brief hypoxic event, whereas Manns et al. examined a group of severely amnesic patients that consisted of 2 cardiac arrest patients, 2 heroin overdose patients, 1 carbon monoxide poisoning patient, and 2 patients with unknown etiologies. We also consider an alternative explanation proposed by Wixted and Squire (2004), who argued that the two patient groups suffered similar deficits, but that statistical or methodological artifacts distorted the results of each of Yonelinas et al.'s experiments. A consideration of those results, however, indicates that such an explanation does not account for the existing data. All of the existing evidence indicates that recollection, but not familiarity, is disrupted in mild hypoxic patients. In more severe cases of hypoxia, or those with more complex etiologies such as heroin overdose, more profound deficits may be observed.

The neural substrates of visual implicit memory: do the two hemispheres play different roles?

Authors:

  • Neal E. A. Kroll

  • Andrew P. Yonelinas

  • Mark M. Kishiyama

  • Kathleen Baynes

  • Robert T. Knight

  • Michael S. Gazzaniga

Date: 2003

PubMed: 14511536

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Abstract:

Identification of visually presented words is facilitated by implicit memory, or visual priming, for past visual experiences with those words. There is disagreement over the neuro-anatomical substrates of this form of implicit memory. Several studies have suggested that this form of priming relies on a visual word-form system localized in the right occipital lobe, whereas other studies have indicated that both hemispheres are equally involved. The discrepancies may be related to the types of priming tasks that have been used because the former studies have relied primarily on word-stem completion tasks and the latter on tasks like word-fragment completion. The present experiments compared word-fragment and word-stem measurements of visual implicit memory in patients with right occipital lobe lesions and patients with complete callosotomies. The patients showed normal visual implicit memory on fragment completion tests, but essentially no visual priming on standard stem completion tests. However, when we used a set of word stems that had only one correct solution for each test item, as was true of the items in the fragment completion tests, the patients showed normal priming effects. The results indicate that visual implicit memory for words is not solely dependent upon the right hemisphere, rather it reflects changes in processing efficiency in bilateral visual regions involved in the initial processing of the items. However, under conditions of high lexical competition (i.e., multiple completion word stems), the lexical processes, which are dominant in the left hemisphere, overshadow the visual priming supported by the left hemisphere.

Effects of extensive temporal lobe damage or mild hypoxia on recollection and familiarity

Authors:

  • Andrew P. Yonelinas

  • Neal E. A. Kroll

  • Joel R. Quamme

  • Michele Lazzara

  • Mary Jane Sauve

  • Keith F. Widaman

  • Robert T. Knight

Date: 2002

PubMed: 12379865

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Abstract:

Memory for past events can be based on recollection or on assessments of familiarity. These two forms of human memory have been studied extensively by philosophers and psychologists, but their neuroanatomical substrates are largely unknown. Here we examined the brain regions that are involved in these two forms of memory by studying patients with damage to different temporal lobe regions. Our results come from (i) structural covariance modeling of recall and recognition, (ii) introspective reports during recognition and (iii) analysis of receiver operating characteristics. In sum, we found that the regions disrupted in mild hypoxia, such as the hippocampus, are centrally involved in conscious recollection, whereas the surrounding temporal lobe supports familiarity-based memory discrimination.

Selective impairment of reasoning about social exchange in a patient with bilateral limbic system damage

Authors:

  • Valerie E. Stone

  • Leda Cosmides

  • John Tooby

  • Neal E. A. Kroll

  • Robert T. Knight

Date: 2002

PubMed: 12177408

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Abstract:

Social exchange is a pervasive feature of human social life. Models in evolutionary biology predict that for social exchange to evolve in a species, individuals must be able to detect cheaters (nonreciprocators). Previous research suggests that humans have a cognitive mechanism specialized for detecting cheaters. Here we provide neurological evidence indicating that social exchange reasoning can be selectively impaired while reasoning about other domains is left intact. The patient, R.M., had extensive bilateral limbic system damage, affecting orbitofrontal cortex, temporal pole, and amygdala. We compared his performance on two types of reasoning problem that were closely matched in form and equally difficult for control subjects: social contract rules (of the form, "If you take the benefit, then you must satisfy the requirement") and precaution rules (of the form, "If you engage in hazardous activity X, then you must take precaution Y"). R.M. performed significantly worse in social contract reasoning than in precaution reasoning, when compared both with normal controls and with other brain-damaged subjects. This dissociation in reasoning performance provides evidence that reasoning about social exchange is a specialized and separable component of human social intelligence, and is consistent with other research indicating that the brain processes information about the social world differently from other types of information.

Visual implicit memory in the left hemisphere: evidence from patients with callosotomies and right occipital lobe lesions

Authors:

  • Andrew P. Yonelinas

  • Neal E. A. Kroll

  • Kathleen Baynes

  • Ian G. Dobbins

  • C.M. Frederick

  • Robert T. Knight

  • Michael S. Gazzaniga

Date: 2001

PubMed: 11476095

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Abstract:

Identification of visually presented objects and words is facilitated by implicit memory for past visual experiences with those items. Several behavioral and neuroimaging studies suggest that this form of memory is dependent on perceptual processes localized in the right occipital lobe. We tested this claim by examining implicit mem- ory in patients with extensive right occipital lobe lesions, using lexi- cal-decision, mirror-reading, picture-fragment, and word-fragment- completion tests, and found that these patients exhibited normal levels of priming. We also examined implicit memory in patients with com- plete callosotomies, using standard and divided-visual-field word- fragment-completion procedures, and found that the isolated left hemisphere exhibited normal priming effects. The results indicate that the right occipital lobe does not play a necessary role in visual im- plicit memory, and that the isolated left hemisphere can support nor- mal levels of visual priming in a variety of tasks.

The contribution of recollection and familiarity to yes-no and forced-choice recognition tests in healthy subjects and amnesics

Authors:

  • Wayne Khoe

  • Neal E. A. Kroll

  • Andrew P. Yonelinas

  • Ian G. Dobbins

  • Robert T. Knight

Date: 2000

PubMed: 10869576

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Abstract:

Recent reports suggest that some amnesic patients perform relatively normally on forced-choice recognition memory tests. Their preserved performance may re ̄ect the fact that the test relies more heavily on assessments of familiarity, a process that is relatively preserved in these patients, than do other recognition tests such as yes±no tests, which may rely more on recollection. The current study examined recognition memory using yes±no and forced-choice procedures in control and amnesic patients in order to determine whether the two tasks di€erentially relied on recollection and familiarity, and whether the extent of the recognition memory de®cit observed in amnesia was dependent upon the type of recognition test used to measure performance. Results using the remember±know procedure with healthy subjects showed that there were no substantial di€erences in recognition accuracy or in the contribution of recollection to these two tasks. Moreover, amnesic patients were not found to perform better on a forced-choice test than on a yes±no test, suggesting that familiarity contributed equally to these two types of recognition test.

The neural substrates of recollection and familiarity (pp. 468-469)

Authors:

  • Andrew P. Yonelinas

  • Neal E. A. Kroll

  • Ian G. Dobbins

  • Michele Lazzara

  • Robert T. Knight

Date: 1999

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Abstract:

Aggleton & Brown argue that a hippocampal-anterior thalamic system supports the “recollection” of contextual information about previ- ous events, and that a separate perirhinal-medial dorsal thalamic system supports detection of stimulus “familiarity.” Although there is a growing body of human literature that is in agreement with these claims, when rec- ollection and familiarity have been examined in amnesics using the process dissociation or the remember/know procedures, the results do not seem to provide consistent support. We reexamine these studies and describe the results of an additional experiment using a receiver operating charac- teristic (ROC) technique. The results of the reanalysis and the ROC ex- periment are consistent with Aggleton & Brown’s proposal. Patients with damage to both regions exhibit severe deficits in recollection and smaller, but consistent, deficits in familiarity.

Recollection and familiarity deficits in amnesia: convergence of remember-know, process dissociation, and receiver operating characteristic data

Authors:

  • Andrew P. Yonelinas

  • Neal E. A. Kroll

  • Ian G. Dobbins

  • Michele Lazzara

  • Robert T. Knight

Date: 1998

PubMed: 9673991

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Abstract:

Previous studies using the process dissociation and the remember-know procedures led to conflicting conclusions regarding the effects of anterograde amnesia on recollection and familiarity. We argue that these apparent contradictions arose because different models were used to interpret the results and because differences in false-alarm rates between groups biased the estimates provided by those models. A reanalysis of those studies with a dual-process signal-detection model that incorporates response bias revealed that amnesia led to a pronounced reduction in recollection and smaller but consistent reduction in familiarity. To test the assumptions of the model and to further assess recognition deficits in amnesics, we examined receiver operating characteristics (ROCs) in amnesics and controls. The ROCs of the controls were curved and asymmetrical, whereas those of the amnesics were curved and symmetrical. The results supported the predictions of the model and indicated that amnesia was associated with deficits in both recollection and familiarity.

Unilateral medial temporal lobe memory impairment: type deficit, function deficit or both?

Authors:

  • Ian G. Dobbins

  • Neal E. A. Kroll

  • Endel Tulving

  • Robert T. Knight

  • Michael S. Gazzaniga

Date: 1997

PubMed: 9539232

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Abstract:

Previous research has characterized memory deficits resulting from unilateral hippocampal system damage as 'material specific', suggesting that left damage results in verbal memory impairment with preservation of visuospatial function and the converse with right damage. Implicit within this hypothesis are the assumptions that the systems are independent and memory is lateralized for each type of material. To test the verbal component of this hypothesis, unilateral hippocampal lesion and commissurotomy patients were compared with controls on a multiple-list free-recall task. The material specific hypothesis predicts severe impairment only with left lesions; right lesions and commissurotomy patients should be only minimally impaired. However, secondary memory was compromised at immediate recall for all patient groups, with both unilateral groups showing comparable and severe verbal episodic memory deficits. Final testing across all lists also revealed severe impairment in commissurotomy patients. Finding both unilateral groups to be similarly impaired for verbal material is taken as evidence against a material specific deficit during this verbal episodic memory task. Although previous data suggest that left patients are considerably more impaired during some verbal tasks, this may not be specific to the material, but rather the combination of material and task demands. Implications for the material specific hypothesis are discussed.

Retrieval of old memories: the temporofrontal hypothesis


Authors:

  • Neal E. A. Kroll

  • Hans J Markowitsch

  • Robert T. Knight

  • D. Yves Von Cramon

Date: 1997

PubMed: 9278629

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Abstract:

Extensive neuropsychological testing is reported on two chronic patients with combined temporopolar and prefrontal damage, dominantly left-hemispheric, and, as control, one chronic patient with bi-hemispheric prefrontal damage. The principal finding is that combined temporofrontal damage, but not substantial prefrontal damage alone, results in marked retrograde memory deficits while leaving intelligence and new learning relatively unimpaired. Although their general world knowledge was good, the temporopolar patients demonstrated retrograde memory impairments on several tests of past events and faces of previously famous people. With respect to retrograde autobiographical memory, one of the temporopolar patients was severely impaired and the other was judged to be moderately impaired. The control patient appeared to be normal. These results, together with corresponding data from related single case and dynamic imaging studies, strongly support a crucial role of the temporofrontal junction area in the ecphory of old memories.

Cohesion failure as a source of memory illusions

Authors:

  • Neal E. A. Kroll

  • Robert T. Knight

  • Janet Metcalfe

  • Elizabeth S. Wolf

  • Endel Tulving

Date: 1996

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Abstract:

One source of ‘‘false’’ memories may be that often only memory fragments are retained. This would then result in a person being unable to distinquish a false conjunction, constructed of memory components, from what had been actually experienced. Experiment 1, employing two- syllable words in a continuous recognition paradigm, found that patients with left hippocampal damage classified more new verbal conjunctions as ‘‘old’’ than did normal subjects or patients with only right hippocampal damage. Experiment 2, employing simple face drawings in a study- test paradigm, found that patients with damage to either side of their hippocampal formation made more conjunction errors with pictorial stimuli than did normal subjects. The results are seen as supporting the hypothesis that binding is an important early step in the consolidation process and that the hippocampal system is a critical component of the neural system involved in the appropriate binding of memory components.